Volume 20

Volume 20 Part 2

Pages
pp. 69 -
90

A brief review of distirbuiton of populations of Rapana venosa in the World Ocean is presented. Short information concerning the formation of metapopulation of R. venosa in the Black Sea and taxonomical history of the species are given. The ecomorphs found off Crimean shores were examined with the emphasis to ecological conditions. It has been demonstrated that presently Rapana metapopulation in the Black Sea consists of numerous populations which differ primarily in conchological characters and size structure. Trophic specialization entails appearance of dwarf forms and influences formation of shell coloration. Studies of protoconchs allowed to conclude that colour pattern is determined on the earliest stages of shell development and later is influenced by environmental factors during the shell growth. The shell characters were compared in several Rapana populations and intrapopulation variability, age and sexual differences were demonstrated. Starting in 2007 the relative shell growth has been increasing despite the normal tendency of slowing of the growth rate with age. This phenomenon is connected with improvement of ecological situation in general, and, the most important, with increasing of forage reserve of Rapana. The pathways of infraspecies forms formation are presented in relation to possible scenarios of environmental changes.

Морфогенез раковины и внутривидовая дифференциация рапаны Rapana venosa (Valenciennes, 1846)
Бондарев И.П.;
Pages
pp. 91 -
101

The shell and radula comparative and illustrated description of all three genera of the family Valvatidae (Valvata s.str., Cincinna and Borysthenia), inhabiting the European fresh-waters are provided. Special emphasis paid to the embryonic shell and radula morphology of Cincinna piscinalis (Müller, 1774) — one of the most common valvatid species in Ukraine. Details of the protoconch of Borysthenia menkeana (Jelski, 1863) is studied with aid of the scanning electron microscopy for the first time. Embryonic shell of those species belonging to all genera discussed possesses identical type of ornamentation («woolen stocking »); it is revealed that valvatid species can be distinguished by proportions of their protoconch and partly by its size. Differences in dimensional characteristics of a protoconch confirm and complete the concept of generic independence of the discussed valvatid groups. Radula of valvatid is highly variable intraspecifically; its aptitude for the species differentiation is limited and requires of additional detailed study of significant set of new data. 

Сравнительная морфология раковины и радулы брюхоногих моллюсков семейства Valvatidae из Северного Причерноморья
Анистратенко О.; Дегтяренко Е.; Анистратенко В.;
Pages
pp. 103 -
108

The new localities of Boreoelona sibirica, at East Siberian snail species (Mollusca: Gastropoda: Bithyniidae), are presented. Descriptions of the shell and copulative apparatus of representatives of the species from waterbodies of Western Siberia and Middle Urals are provided.

Boreoelona sibirica Westerlund, 1886) (Gastropoda, Pectinibranchia, Bithynidae) в водоемах Западной Сибири и Среднего Урала.
Лазуткина Е.А.; Андреева С.И.; Андреев Н.И. ;
Pages
pp. 109 -
115

Illustrated descriptions of Columella talgarica sp. nov., Leucozonella corona sp. nov., and Angiomphalia nucula sp. nov. from Kazakhstan and Kirgisia are given. Holotypes of the species are deposited in Zoological Museum of Moscow State University (ЗММУ, ZMMU).

Три новых вида наземных моллюсков (Pulmonata: Truncatellinidae et Hygromiidae) из Центральной Азии
Шилейко А.А.; Рымжанов Т.С.;
Pages
pp. 117 -
139

On the basis of the molecular sequences of 12S r-RNA gene of 16 species of Costellariidae the phylogenetic hypothesis is proposed. The anatomy of the foregut and radula of 15 species of the family was studied. The anterior foregut is the most rich in phylogenetically informative characters in Costellariidae and Neogastropoda in general. 6 types of anatomy of the mid-oesophagus have been recognized. They differ in the anatomy of the gland of Leiblein and in the presence/absence and state of development of the glandular tube, formed by stripping off and closure of the glandular folds of mid-oesophagus. On the basis of molecular phylogeny it was demonstrated that the most basal underived type of anterior foregut was found in still unnamed species of the family. It is characterized by unmodified gland of Leiblein opening directly into glandular mid-oesophagus. It is proved that the process of the formation of the glandular tube and transformation of the gland of Leiblein into muscular organ occurred in Costellariidae independently from other lineages of Neogastropoda. The functional morphology of the digestive system is discussed. The most probable source of toxins in costellariids is despite expectations not the gland of Leiblein, but accessory or primary salivary glands.

Эволюция хищных брюхоногих моллюсков семейства Costellariidae (Neogastropoda) в рамках молекулярной филогении
Федосов А.Э.; Кантор Ю.И.;